1. Undergraduate Education:
Calvin College, Grand Rapids, Michigan, B.A. in Biology with honors, 09/1979 – 05/1983
2. Graduate Education:
Case Western Reserve University, Cleveland, OH, Ph.D. in Molecular Biology and Microbiology, 06/1985 – 01/1991, Thesis: “Modulation of CSV-5’ and 3’ LTR transcription in c-myc activation”, Advisor: Dr. H.-J. Kung
Case Western Reserve University School of Medicine, Cleveland, OH, M.D., 08/1983 – 06/1985; 01/1991 – 01/1993
3. Postgraduate Training:
National Institutes of Health, NIAMS, Bethesda, MD, “Acid maltase deficiency”, 05/1992 – 06/1992, 01/1993 – 06/1993, Mentor: Dr. Paul Plotz
University of Washington, Seattle, WA, Pediatric Residency Program (PGY-1, PGY-2, PGY-3), 06/1993 – 06/1996
The Hospital for Sick Children, Toronto, ON, Clinical Genetics Fellowship, 07/1996 – 06/1998
Baylor College of Medicine, Houston, TX, “Murine model of Smith-Magenis Syndrome and Charcot-Marie-Tooth Disease”, 07/1998 – 09/2001, Mentor: Dr. James Lupski.
Baylor College of Medicine, Houston, TX, “Fly and mouse models of Schimke immunoosseous dysplasia”, 09/2001 – 10/2002, Mentor: Dr. Hugo Bellen.